Kenneth R. Robertson, Loy R. Phillippe, Geoffey A. Levin, and Michael J. Moore
Center for Biodiversity
Illinois Natural History Survey
607 East Peabody Drive
Champaign, IL 61820 USA
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In 1917, at the height of World War I, the United States Army was searching for remote, sparsely populated sites to test and store munitions. A huge expanse of sand near the Mississippi River in northwestern Illinois (Carroll and Jo Daviess counties) fit the bill and the Savanna Army Depot was established. Prior to the Army's occupation, this area, known locally as "The Prairie," was studied and observed by Dr. Henry Allan Gleason in 1908. Now, 80 years later, the Depot is being decommissioned. The Illinois Department of Natural Resources, together with various federal agencies, is gathering data to help plan the Depot's future. A team of botanists from the Illinois Natural History Survey's Center for Biodiversity is responsible for evaluating the natural communities and surveying the plants that exist at the Depot.
The Army probably did not recognize the tremendous biological significance of the Depot when it was established, but its extensive sands are very unusual in Illinois. Previous DNR-sponsored surveys of the Depot had located 11 threatened or endangered plant species. With increased access to the base, we discovered one additional state endangered species and many previously unknown populations of the others. We also found at least one plant previously unknown from Illinois that should be a candidate for threatened or endangered status.
Many of the Depot's rare plants are western species that reach their eastern limits on sandy habitats in western Illinois. Two of these, Mirabilis hirsuta (hairy umbrella wort) and Opuntia fragilis (fragile prickly pear), are known in Illinois only in Jo Daviess County, and the latter is known only from the Savanna Army Depot. Others, like Polanisia jamesii (James' clammyweed) and Carex tonsa (shaved sedge) have their largest Illinois populations on the Depot. Some of the Depot's animals show a similar pattern. For example, the only population of Lepus townsendii (white-tailed jackrabbits) in Illinois was on the Depot. Unfortunately, the jackrabbits disappeared in recent years, probably due to habitat changes and increased numbers of coyotes.
Perhaps even more significant than the individual plant species are the Depot's natural communities. Most noteworthy are its extensive prairies, savannas, and river dune forest. The savanna habitat, which consists of trees, typically oaks, scattered through a grassland, has almost disappeared from North America. Only about 6,400 acres (2,591 hectares) of tallgrass savanna remain in the United States, about 0.02% of the presettlement total. Prairies have seen similar losses. The Depot, with about 4,500 acres (1,822 hectares) of savanna and prairie, has the most extensive remaining stands of these communities in the state.
The natural communities of the Depot are not pristine, however. For obvious reasons, the Army has vigorously suppressed fires, which are vital for preserving both prairie and savanna from encroaching forests. Cattle have been used to reduce grassy vegetation and therefore reduce fire intensity. Though the cattle probably have slowed forest encroachment, they have also modified the grassland and forest vegetation. We have been uniquely able to document changes in the Depot's vegetation because not long before the Army acquired the land, noted ecologist Henry Allan Gleason, then at the Natural History Survey, published a paper describing the vegetation of the major sand deposits in Illinois, including the region that became the Depot. Overall, we observed that forest cover of the depot has increased, particular in areas that formerly were savanna. In addition, the abundance of some shrubs and grasses has increased, whereas other late successional grasses and forbs that are probably dependent on periodic fires have decreased in abundance.
Still, much of the Savanna Army Depot remains in vegetation that with proper management could recover significantly. Nowhere else in Illinois can one look across a landscape so extensively covered with prairie and savanna. As decisions are being made for the Depot's future, our research provides useful information will help preserve this remarkable remnant of our state's natural heritage.
A total of 617 taxa (species, subspecies, varieties, and forms) were found during 1996. Gleason (1910) recorded several other plants that we did not observe in 1996, including: Smilacina racemosa, Smilax ecirrhata, Petalostemum candidum, Asclepias exaltata, Scutellaria parvula / S. leonardii, Liatris cylindracea, Solidago missouriensis, and Parthenium integrifolium. Of the total of 617 taxa, 480 are native species, 3 are questionably native, and 134 are introduced.
Below is an annotated List of Vascular Plants at the Savanna Army Depot. Arranged alphabetically by scientific name. This list includes only species we observed during 1996, except for Polygala verticillata, which was collected in 1993 and Orobanche fasciculata (Orobanchaceae), which was last collected at the Depot in 1908 although suitable habitat is still extant. Under "Scientific Names", the authorities are omitted for brevity; they are included on the voucher specimens cited in Table 2. Under "E/T", E = State Endangered, T = State Threatened, FCC = former Federal Candidate Category 2, ND = Newly Discovered in Illinois and likely to qualify for listing as a state endangered species.
Scientific Name
|
Common Name
|
Family
|
Native or Introduced
|
Frequency
and Habitat
|
E/T
|
| Acalypha
rhomboidea
|
Three-seeded
Mercury
|
Euphorbiaceae
|
native
|
Occasional
in sand forest and in wet-mesic floodplain forest
|
--
|
| Acer
negundo
|
Boxelder
|
Aceraceae
|
native
|
Occasional
in sand forest, sand savanna, pastureland, and wet-mesic floodplain forest;
infrequent in sand prairie
|
--
|
| Acer
saccharinum
|
Silver
maple
|
Aceraceae
|
native
|
Abundant
in wet-mesic floodplain forest
|
--
|
| Acer
saccharum
|
Sugar
Maple
|
Aceraceae
|
native
|
Occasional
in sand forest
|
--
|
| Achillea
millefolium
|
Yarrow
|
Asteraceae
|
introduced
|
Frequent
in sand prairie and sand savanna; occasional in pastureland
|
--
|
| Adiantum
pedatum
|
Maidenhair
Fern
|
Adiantaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Agalinis
tenuifolia
|
Common
Agalinis
|
Scrophulariaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Agastache
nepetoides
|
Yellow
Giant Hyssop
|
Lamiaceae
|
native
|
Occasional
in sand savanna
|
--
|
| Agrimonia
gryposepala
|
Tall
Agrimony
|
Rosaceae
|
native
|
Occasional
in sand forest and sand savanna
|
--
|
| Agrimonia
pubescens
|
Soft
Agrimony
|
Rosaceae
|
native
|
Occasional
in sand forest
|
--
|
| Agropyron
repens f. aristatum
|
Quack
Grass
|
Poaceae
|
introduced
|
Occasional
in sand prairie
|
--
|
| Agropyron
repens f. repens
|
Quack
Grass
|
Poaceae
|
introduced
|
Occasional
in sand prairie
|
--
|
| Agropyron
smithii
|
Western
Wheat Grass
|
Poaceae
|
introduced
|
Occasional
in developed land, sand prairie and sand savanna
|
--
|
| Agropyron
subsecundum
|
Bearded
Wheat Grass
|
Poaceae
|
native
|
Infrequent
in sand savanna
|
E
|
| Agrostis
gigantea
|
Redtop
|
Poaceae
|
introduced
|
Infrequent
in developed land and sand prairie
|
--
|
| Agrostis
hyemalis
|
Bent
Grass
|
Poaceae
|
native
|
Occasional
in developed land, sand prairie, and sand savanna
|
--
|
| Alisma
plantago-aquatica var. parviflorum
|
Small-flowered
Water Plantain
|
Alismataceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Alliaria
petiolata
|
Garlic
Mustard
|
Brassicaceae
|
introduced
|
Frequent
to abundant in sand forest
|
--
|
| Allium
canadense
|
Wild
Onion
|
Liliaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Alopecurus
carolinianus
|
Common
Foxtail
|
Poaceae
|
native
|
Occasional
in disturbed sand pond
|
--
|
| Alyssum
alyssoides
|
Pale
Allysum
|
Brassicaceae
|
introduced
|
Occasional
in developed land and sand prairie
|
--
|
| Amaranthus
rudis
|
Water
Hemp
|
Amaranthaceae
|
native
|
Frequent
in wet-mesic floodplain forest
|
--
|
| Amaranthus
spinosus
|
Spiny
Amaranth
|
Amaranthaceae
|
introduced
|
Rare
in wet-mesic floodplain forest
|
--
|
| Amaranthus
tuberculatus
|
Amaranth
|
Amaranthaceae
|
introduced
|
Frequent
in wet-mesic floodplain forest
|
--
|
| Ambrosia
artemisiifolia
|
Common
Ragweed
|
Asteraceae
|
native?
|
Infrequent
in developed land and sand prairie
|
--
|
| Ambrosia
psilostachya
|
Western
Ragweed
|
Asteraceae
|
native
|
Common
in sand prairie
|
|
| Ambrosia
trifida
|
Giant
Ragweed
|
Asteraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Ammannia
coccinea
|
Tooth-cup
|
Lythraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Amorpha
canescens
|
Leadplant
|
Fabaceae
|
native
|
Occasional
in sand prairie
|
--
|
| Amorpha
fruticosa
|
False
Indigo
|
Fabaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Amphicarpa
bracteata
|
Hog-peanut
|
Fabaceae
|
native
|
Occasional
in wet-mesic floodplain forest; infrequent in sand forest
|
--
|
| Andropogon
gerardii
|
Big
Bluestem
|
Poaceae
|
native
|
Occasional
in sand prairie
|
--
|
| Androsace
occidentalis
|
Rock
Jasmine
|
Primulaceae
|
native
|
Frequent
in sand prairie and sand savanna
|
--
|
| Anemone
canadensis
|
Meadow
Anemone
|
Ranunculaceae
|
native
|
Frequent
in border with sand forest and wet-mesic floodplain forest
|
--
|
| Anemone
caroliniana
|
Carolina
Anemone
|
Ranunculaceae
|
native
|
Infrequent
in sand prairie
|
--
|
| Anemone
cylindrica
|
Thimbleweed
|
Ranunculaceae
|
native
|
Occasional
in sand forest
|
--
|
| Anemone
quinquefolia
|
Wood
Anemone
|
Ranunculaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Anemone
virginiana
|
Tall
Anemone
|
Ranunculaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Antennaria
neglecta
|
Cat's
Foot
|
Asteraceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Antennaria
plantaginifolia
|
Pussy
Toes
|
Asteraceae
|
native
|
Infrequent
in sand savanna
|
--
|
| Apios
americana
|
Common
Ground-nut
|
Fabaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Apocynum
sibiricum
|
Creeping
Dogbane
|
Apocynaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Aquilegia
canadensis
|
Wild
Columbine
|
Ranunculaceae
|
native
|
Occasional
in sand forest
|
--
|
| Arabis
canadensis
|
Sicklepod
|
Brassicaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Arabis
glabra
|
Tower
Mustard
|
Brassicaceae
|
native
|
Rare
in sand prairie
|
--
|
| Arabis
lyrata
|
Sand
Cress
|
Brassicaceae
|
native
|
Common
in sand prairie and sand savanna
|
--
|
| Arabis
x divaricarpa
|
Purple
Rock Cress
|
Brassicaceae
|
native
|
Infrequent
in sand prairie
|
ND
|
| Aralia
nudicaulis
|
Wild
Sarsaparilla
|
Araliaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Arctium
lappa
|
Great
Burdock
|
Asteraceae
|
introduced
|
Infrequent
in sand savanna
|
--
|
| Arctium
minus
|
Common
Burdock
|
Asteraceae
|
introduced
|
Occasional
in disturbed sand savanna
|
--
|
| Arenaria
serpyllifolia
|
Thyme-leaved
Sandwort
|
Caryophyllaceae
|
introduced
|
Frequent
in developed land and occasional in sand prairie
|
--
|
| Arisaema
dracontium
|
Green
Dragon
|
Araceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Arisaema
triphyllum
|
Jack-in-the-Pulpit
|
Araceae
|
native
|
Infrequent
in sand forest
|
--
|
| Aristida
basiramea
|
Forktip
Three-awn Grass
|
Poaceae
|
native
|
Frequent
in sand prairie and sand savanna
|
--
|
| Aristida
oligantha
|
Prairie
Three-awn Grass
|
Poaceae
|
native
|
Infrequent
in sand prairie and successional field
|
--
|
| Aristida
tuberculosa
|
Beach
Three-awn Grass
|
Poaceae
|
native
|
Frequent
in sand prairie and sand savana; Infrequent in sand forest
|
--
|
| Artemisia
campestris
|
Wormwood
|
Asteraceae
|
native
|
Occasional
in sand forest, sand prairie, and sand savanna
|
--
|
| Artemisia
ludoviciana
|
White
Sage
|
Asteraceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Asclepias
amplexicaulis
|
Sand
Milkweed
|
Asclepiadaceae
|
native
|
infrequent
in sand prairie
|
--
|
| Asclepias
incarnata
|
Swamp
Milkweed
|
Asclepiadaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Asclepias
syriaca
|
Common
Milkweed
|
Asclepiadaceae
|
native
|
Infrequent
in successional field
|
--
|
| Asclepias
tuberosa
|
Butterflyweed
|
Asclepiadaceae
|
native
|
Rare
in sand savanna
|
--
|
| Asclepias
verticillata
|
Horsetail
Milkweed
|
Asclepiadaceae
|
native
|
Frequent
in sand prairie; occasional in sand savanna; infrequent in pastureland
|
--
|
| Asclepias
viridiflora
|
Tall
Green Milkweed
|
Asclepiadaceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Asparagus
officinalis
|
Asparagus
|
Liliaceae
|
introduced
|
Occasional
in sand savanna
|
--
|
| Asplenium
platyneuron
|
Ebony
Spleenwort
|
Aspleniaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Aster
cordifolius
|
Heart-leaved
Aster
|
Asteraceae
|
native
|
Infrequent
in sand forest
|
--
|
| Aster
ericoides
|
White
Prairie Aster
|
Asteraceae
|
native
|
Occasional
in sand prairie
|
--
|
| Aster
lanceolatus
|
Panicled
Aster
|
Asteraceae
|
native
|
Frequent
in wet-mesic floodplain forest
|
--
|
| Aster
lateriflorus
|
Starved
Aster
|
Asteraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Aster
linariifolius
|
Flax-leaved
Aster
|
Asteraceae
|
native
|
Occasional
in sand forest, sand prairie, and sand savana
|
--
|
| Aster
oblongifolius
|
Aromatic
Aster
|
Asteraceae
|
native
|
Infrequent
in sand forest and sand prairie
|
--
|
| Aster
ontarionis
|
Ontario
Aster
|
Asteraceae
|
native
|
Frequent
in wet-mesic floodplain forest
|
--
|
| Aster
oolentangiensis
|
Sky
Blue Aster
|
Asteraceae
|
native
|
Rare
in sand savanna
|
--
|
| Aster
pilosus
|
Hairy
Aster
|
Asteraceae
|
native
|
Common
in sand prairie and sand savanna
|
|
| Aster
prenanthoides
|
Crooked-stemmed
Aster
|
Asteraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Aster
puniceus
|
Purple-stemmed
Aster
|
Asteraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Aster
sericeus
|
Western
Silvery Aster
|
Asteraceae
|
native
|
Infrequent
in sand prairie
|
--
|
| Athyrium
filix-femina var. angustum
|
Lady
Fern
|
Dryopteridaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Bacopa
rotundifolia
|
Water
Hyssop
|
Scrophulariaceae
|
native
|
Rare
in natural sand pond
|
--
|
| Barbarea
vulgaris
|
Yellow
Rocket
|
Brassicaceae
|
introduced
|
Infrequent
in developed land
|
--
|
| Berberis
thunbergii
|
Japanese
Barberry
|
Berberidaceae
|
introduced
|
Infrequent
in sand forest and sand savanna
|
--
|
| Berteroa
incana
|
Hoary
Alyssum
|
Brassicaceae
|
introduced
|
Occasional
in developed land, sand prairie and sand savanna
|
--
|
| Besseya
bullii
|
Kitten
Tails
|
Scrophulariaceae
|
native
|
Occasional
in sand forest, sand prairie, and sand savanna
|
T
|
| Betula
nigra
|
River
Birch
|
Betulaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Bidens
cernua
|
Bur-marigold
|
Asteraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Bidens
tripartita
|
Beggar-ticks
|
Asteraceae
|
introduced
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Bidens
vulgata
|
Tall
Beggar-ticks
|
Asteraceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Boehmeria
cylindrica
|
False
Nettle
|
Urticaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Botrychium
dissectum
|
Dissected
Grapefern
|
Ophioglossaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Botrychium
virginianum
|
Rattlesnake
Fern
|
Ophioglossaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Bouteloua
curtipendula
|
Side-oats
Grama Grass
|
Poaceae
|
native
|
Occasional
in sand prairie
|
--
|
| Bouteloua
gracilis
|
Blue
Grama-grass
|
Poaceae
|
native
|
Rare
in sand prairie
|
ND
|
| Bouteloua
hirsuta
|
Hairy
Grama Grass
|
Poaceae
|
native
|
Frequent
in sand prairie
|
--
|
| Brassica
nigra
|
Black
Mustard
|
Brassicaceae
|
introduced
|
Infrequent
in developed land
|
|
| Brickellia
eupatorioides
|
False
Boneset
|
Asteraceae
|
native
|
Infrequent
in sand prairie
|
--
|
| Bromus
inermis
|
Awnless
Brome
|
Poaceae
|
introduced
|
Occasional
in developed land and sand prairie
|
--
|
| Bromus
kalmii
|
Kalm's
Brome
|
Poaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Bromus
racemosus
|
Smooth
Chess
|
Poaceae
|
introduced
|
Occasional
to frequent in sand prairie and sand savanna
|
--
|
| Bromus
tectorum
|
Downy
Chess
|
Poaceae
|
introduced
|
Frequent
in developed land and sand prairie and sand savanna
|
--
|
| Bulbostylis
capillaris
|
Autumn
Sedge
|
Cyperaceae
|
native
|
Occasional
in sand prairie
|
--
|
| Calamovilfa
longifolia
|
Sand-reed
|
Poaceae
|
native
|
Rare
in sand savanna
|
--
|
| Callirhoë
triangulata
|
Poppy
Mallow
|
Malvaceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Caltha
palustris
|
Marsh
Marigold
|
Ranunculaceae
|
native
|
Rare
in wet-mesic floodplain forest
|
--
|
| Calystegia
sepium
|
American
Bindweed
|
Convolvulaceae
|
native
|
Infrequent
in developed land along edge of wet-mesic floodplain forest
|
--
|
| Campanula
americana
|
American
Bellflower
|
Campanulaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Cannabis
sativa
|
Marijuana
|
Cannabinaceae
|
introduced
|
Occasional
in sand savanna
|
--
|
| Capsella
bursa-pastoris
|
Shepard's
Purse
|
Brassicaceae
|
introduced
|
Occasional
in developed land
|
--
|
| Cardamine
bulbosa
|
Spring
Cress
|
Brassicaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Cardamine
hirsuta
|
Spring
Cress
|
Brassicaceae
|
introduced
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Cardamine
parviflora var. arenicola
|
Small-flowered
Bitter Cress
|
Brassicaceae
|
native
|
Occasional
in sand savanna
|
--
|
| Cardamine
pensylvanica
|
Pennsylvania
Bitter-cress
|
Brassicaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Carduus
nutans
|
Musk
Thistle
|
Asteraceae
|
introduced
|
Infrequent
in successional field
|
--
|
| Carex
annectens
|
Large
Yellow Fox Sedge
|
Cyperaceae
|
native
|
Occasional
in sand prairie, edge of ponds, and wet-mesic floodplain forest
|
--
|
| Carex
bicknellii
|
Copper-shouldered
Oval Sedge
|
Cyperaceae
|
native
|
Frequent
in sand prairie and sand savanna
|
--
|
| Carex
blanda
|
Common
Wood Sedge
|
Cyperaceae
|
native
|
Occasional
in sand forest
|
--
|
| Carex
brevior
|
Plains
Oval Sedge
|
Cyperaceae
|
native
|
Frequent
in sand prairie
|
--
|
| Carex
conjuncta
|
Green-headed
Fox Sedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Carex
cristatella
|
Crested
Oval Sedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Carex
duriuscula
|
Sedge
|
Cyperaceae
|
native
|
Rare
in developed land
|
--
|
| Carex
frankii
|
Bristly
Cattail Sedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic bottomland forest and along streams
|
--
|
| Carex
gravida
|
Long-awned
Bracted Sedge
|
Cyperaceae
|
native
|
Occasional
in developed land and sand prairie
|
--
|
| Carex
grayi
|
Common
Bur Sedge
|
Cyperaceae
|
native
|
Occasional
in wet mesic floodplain forest
|
--
|
| Carex
grisea
|
Wood
Gray Sedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Carex
hirtifolia
|
Hairy
Wood Sedge
|
Cyperaceae
|
native
|
Occasional
in sand forest
|
--
|
| Carex
hystericina
|
Porcupine
Sedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Carex
laeviconica
|
Long-toothed
Lake Sedge
|
Cyperaceae
|
native
|
Rare
along edge of ox-bow lake
|
--
|
| Carex
lupulina
|
Common
Hop Sedge
|
Cyperaceae
|
native
|
Occasional
in edge of pond and wet-mesic floodplain forest
|
--
|
| Carex
meadii
|
Mead's
Stiff Sedge
|
Cyperaceae
|
native
|
Infrequent
in sand prairie and sand savanna
|
--
|
| Carex
molesta
|
Field
Oval Sedge
|
Cyperaceae
|
native
|
Occasional
in sand prairie
|
--
|
| Carex
muhlenbergii
|
Sand
Bracted Sedge
|
Cyperaceae
|
native
|
Frequent
in sand prairie and sand savanna; occasional in sand forest
|
--
|
| Carex
pensylvanica
|
Common
Oak Sedge
|
Cyperaceae
|
native
|
Occasional
to frequent in sand forest, sand prairie and sand savanna
|
--
|
| Carex
rosea
|
Curly-styled
Wood Sedge
|
Cyperaceae
|
native
|
Occasional
in sand forest
|
--
|
| Carex
scoparia var. scoparia
|
Lance-fruited
Oval Sedge
|
Cyperaceae
|
native
|
Occasional
in sand savanna
|
--
|
| Carex
stipata
|
Common
Fox Sedge
|
Cyperaceae
|
native
|
Occasional
in sand forest
|
--
|
| Carex
stricta
|
Common
Tussock Sedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Carex
tonsa
|
Shaved
Sedge
|
Cyperaceae
|
native
|
Frequent
in sand prairie and sand savanna; occasional in sand forest
|
E
|
| Carex
tribuloides
|
Awl-fruited
Oval Sedge
|
Cyperaceae
|
native
|
Occasional
in edge of pond
|
--
|
| Carex
typhina
|
Common
Cattail Sedge
|
Cyperaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Carex
vulpinoidea
|
Brown
Fox Sedge
|
Cyperaceae
|
native
|
Occasional
in developed land
|
--
|
| Carya
cordiformis
|
Bitternut
Hickory
|
Juglandaceae
|
native
|
Occasional
in sand forest and wet-mesic floodplain forest
|
--
|
| Carya
ovata
|
Shagbark
Hickory
|
Juglandaceae
|
native
|
Occasional
in sand forest
|
--
|
| Carya
tomentosa
|
Mockernut
Hickory
|
Juglandaceae
|
native
|
Occasional
in sand forest
|
--
|
| Catalpa
speciosa
|
Northern
Catalpa
|
Bignoniaceae
|
introduced
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Ceanothus
americanus
|
New
Jersey Tea
|
Rhamnaceae
|
native
|
Infrequent
in sand savanna
|
--
|
| Ceanothus
herbaceus (C. ovatus)
|
Redroot
|
Rhamnaceae
|
native
|
Occasional
to frequent in sand prairie
|
E
|
| Celastrus
scandens
|
Bittersweet
|
Celastraceae
|
native
|
Occasional
in sand forest
|
--
|
| Celtis
occidentalis
|
Hackberry
|
Ulmaceae
|
native
|
Occasional
in sand forest and sand savanna Infrequent in sand prairie
|
--
|
| Cenchrus
longispinus
|
Sand
Bur
|
Poaceae
|
native
|
Frequent
in developed land and occasional in sand prairie
|
--
|
| Centaurea
maculosa
|
Spotted
Knapweed
|
Asteraceae
|
introduced
|
Occasional
in successional field
|
--
|
| Cephalanthus
occidentalis
|
Buttonbush
|
Rubiaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Cerastium
brachypodum
|
Mouse-eared
Chickweed
|
Caryophyllaceae
|
introduced
|
Occasional
in sand prairie and developed land
|
--
|
| Cerastium
vulgatum
|
Common
Mouse-eared Chickweed
|
Caryophyllaceae
|
introduced
|
Occasional
in developed land and sand prairie and sand savanna
|
--
|
| Ceratophyllum
demersum
|
Hornwort
|
Ceratophyllaceae
|
native
|
Occasional
as aquatic in Mississippi River
|
--
|
| Chaenorrhinum
minus
|
Dwarf
Snapdragon
|
Scrophulariaceae
|
introduced
|
Occasional
in developed land
|
--
|
| Chamaecrista
fasciculata
|
Partridge
Pea
|
Fabaceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Chamaesyce
geyeri
|
Geyer's
Spurge
|
Euphorbiaceae
|
native
|
Frequent
in sand prairie and sand savanna
|
--
|
| Chamaesyce
maculata
|
Spotted
Spurge
|
Euphorbiaceae
|
native
|
Occasional
in developed land
|
--
|
| Chamaesyce
nutans
|
Eyebane
|
Euphorbiaceae
|
native
|
Occasional
in developed land
|
--
|
| Chenopodium
ambrosioides
|
Mexican
Tea
|
Chenopodiaceae
|
introduced
|
Occasional
in developed land and successional field
|
--
|
| Chenopodium
gigantospermum
|
Maple-leaved
Goose Foot
|
Chenopodiaceae
|
native
|
Infrequent
in sand savanna
|
--
|
| Chenopodium
pratericola
|
Goosefoot
|
Chenopodiaceae
|
native
|
Occasional
in sand prairie, sand savanna and sand forest; infrequent in developed land
|
--
|
| Chenopodium
simplex
|
Maple-leaved
Goosefoot
|
Chenopodiaceae
|
native
|
Occasional
in sand forest and sand savanna
|
--
|
| Chloris
verticillata
|
Windmill
Grass
|
Poaceae
|
introduced
|
Occasional
in developed land and successional field
|
--
|
| Cicuta
maculata
|
Water
Hemlock
|
Apiaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Cinna
arundinacea
|
Common
Woodreeed
|
Poaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Circaea
lutetiana subsp. canadensis
|
Enchanter's
Nightshade
|
Onagraceae
|
native
|
Occasional
in sand forest
|
--
|
| Cirsium
arvense
|
Canada
Thistle
|
Asteraceae
|
introduced
|
Infrequent
in pastureland
|
--
|
| Cirsium
discolor
|
Field
Thistle
|
Asteraceae
|
native
|
Infrequent
in wet-mesic bottomland forest
|
--
|
| Cirsium
vulgare
|
Bull
Thistle
|
Asteraceae
|
introduced
|
Infrequent
in sand savanna
|
--
|
| Claytonia
virginica
|
Spring
Beauty
|
Portulacaceae
|
native
|
Frequent
in sand forest
|
--
|
| Clematis
virginiana
|
Virgin's
Bower
|
Ranunculaceae
|
native
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Comandra
umbellata
|
False
Toadflax
|
Santalaceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Commelina
erecta
|
Erect
Day-flower
|
Commelinaceae
|
native
|
Occasional
in sand forest and sand savanna
|
--
|
| Conium
maculatum
|
Poison
Hemlock
|
Apiaceae
|
introduced
|
Infrequent
in wet-mesic floodplain forest
|
--
|
| Convolvulus
arvensis
|
Field
Bindweed
|
Convolvulaceae
|
introduced
|
Occasional
in sand prairie
|
--
|
| Conyza
canadensis
|
Horseweed
|
Asteraceae
|
native
|
Frequent
in successional field, infrequent in sand savanna
|
--
|
| Coreopsis
palmata
|
Prairie
Coreopsis
|
Asteraceae
|
native
|
Infrequent
in sand prairie and sand savanna
|
--
|
| Cornus
drummondii
|
Rough-leaved
Dogwood
|
Cornaceae
|
native
|
Occasional
in sand forest and sand savanna
|
--
|
| Cornus
racemosa
|
Gray
Dogwood
|
Cornaceae
|
native
|
Frequent
in sand forest, sand savanna, and wet-mesic floodplain forest
|
--
|
| Coronilla
varia
|
Crown
Vetch
|
Fabaceae
|
introduced
|
Occasional
in developed land, sand prairie, and sand savanna
|
--
|
| Corydalis
micrantha
|
Slender
Corydalis
|
Fumariaceae
|
native
|
Occasional
in sand forest and sand savanna
|
--
|
| Corylus
americana
|
Hazelnut
|
Betulaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Crataegus
calpodendron
|
Hawthorn
|
Rosaceae
|
native
|
Occasional
in sand forest, sand prairie, and sand savanna
|
--
|
| Crepis
tectorum
|
Hawksbeard
|
Asteraceae
|
introduced
|
Infrequent
in sand prairie and successional field
|
--
|
| Crotalaria
sagittalis
|
Rattlebox
|
Fabaceae
|
native
|
Occasional
in sand prairie and sand savanna
|
--
|
| Croton
glandulosus var. septentrionalis
|
Sand
Croton
|
Euphorbiaceae
|
native?
|
Occasional
in sand prairie and sand savanna
|
--
|
| Cryptotaenia
canadensis
|
Honewort
|
Apiaceae
|
native
|
Abundant
in wet-mesic floodplain forest; occasional in sand forest
|
--
|
| Cycloloma
atriplicifolium
|
Winged
Pigweed
|
Chenopodiaceae
|
native
|
Occasional
in developed land, sand forest, sand prairie, and sand savanna
|
|
| Cynoglossum
officinale
|
Common
Hound's-tongue
|
Boraginaceae
|
introduced
|
Occasiona
in developed land and sand savanna
|
--
|
| Cyperus
aristatus
|
Flatsedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Cyperus
erythrorhizos
|
Redroot
Flatsedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Cyperus
esculentus var. esculentus
|
Yellow
Nutsedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic bottomland forest
|
--
|
| Cyperus
filiculmis var. filiculmis
|
Slender
Sand Sedge
|
Cyperaceae
|
native
|
Frequent
in sand prairie and sand savnaan; occasional in sand forest
|
--
|
| Cyperus
filiculmis var. macilentus
|
Slender
Sand Sedge
|
Cyperaceae
|
native
|
Occasional
in sand forest; frequent in sand prairie and sand savanna
|
--
|
| Cyperus
grayioides
|
Gray's
Umbrella Sedge
|
Cyperaceae
|
native
|
Occasional
in blowouts of sand prairie and sand savanna
|
T
|
| Cyperus
odoratus
|
Flatsedge
|
Cyperaceae
|
native
|
Occasional
in wet-mesic floodplain forest
|
--
|
| Cyperus
schweinitzii
|
Rough
Sand Sedge
|
Cyperaceae
|
native
|
Occasional
in sand forest; frequent in sand prairie and sand savanna
|
--
|
| Cystopteris
protrusa
|
Fragile
Fern
|
Dryopteridaceae
|
native
|
Occasional
in sand forest
|
--
|
| Dactylis
glomerata
|
Orchard
Grass
|
Poaceae
|
introduced
|
Occasional
in developed land
|
--
|
| Daucus
carota
|
Queen
Anne's Lace
|
Apiaceae
|
introduced
|
Occasional
in successional field
|
--
|
| Descurainia
pinnata subsp. brachycarpa
|
Tansy
Mustard
|
Brassicaceae
|
native
|
Infrequent
in sand prairie, sand savanna, and developed land
|
--
|
| Desmodium
canadense
|
Canadian
Tick-trefoil
|
Fabaceae
|
native
|
Infrequent
in opening in wet-mesic floodplain forest
|
--
|
| Desmodium
glutinosum
|
Pointed
Tick Trefoil
|
Fabaceae
|
native
|
Occasional
in sand forest
|
--
|
| Desmodium
illinoense
|
Illinois
Tick Trefoil
|
Fabaceae
|
native
|
Infrequent
in sand prairie
|
--
|
| Dianthus
armeria
|
Deptford
Pink
|
Caryophyllaceae
|
introduced
|
Occasional
in sand prairie and sand savanna
|
--
|
| Dicentra
cucullaria
|
Dutchman's
Breeches
|
Fumariaceae
|
native
|
Infrequent
in sand forest
|
--
|
| Digitaria
sanguinalis
|
Hairy
Crabgrass
|
Poaceae
|
introduced
|
Occasional
in developed land
|
--
|
| Diodia
teres
|
Rough
Buttonweed
|
Rubiaceae
|
native?
|
Frequent
in sand prairie and sand savanna and occasional in sand forest
|
--
|
| Draba
nemorosa var. leiocarpa
|
Whitlow
Grass
|
Brassicaceae
|
native
|
Rare
in sand prairie
|
ND
|
| Draba
reptans
|
Common
Whitlow Grass
|
Brassicaceae
|
native
|
Frequent
in sand prairie and sand savanna
|
--
|
| Draba
verna
|
Vernal
Whitlow Grass
|
Brassicaceae
|
introduced
|
Occasional
in developed land
|
--
|
| Dryopteris
carthusiana
|
Spinulose
Woodfern
|
Dryopteridaceae
|
native
|
Occasional
in sand forest
|
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Updated 8 October 1997